epiEnhancer Atlas

Genetic context, enhancer non-additivity and functional regulatory classes

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Genes

GeneSymbolChrCross-supportInteraction score Best FDRTier 1 pairsoverlapEpiGene setsoverlapEpiEnh sets Overlap enh.Overlap LD blocksExact Tier 1

Enhancers

EnhancerChrCross-supportInteraction genesInteraction pairs Tier 1 pairsOverlap genesOverlap traitsOverlap LD blocksExact genes

FANTOM5 enhancer functional classes

Final 16-subcluster GTM/k-means solution with tissue activity, GO Biological Process and HOMER motif enrichment.

Tissue-class activity

Tissue classActive fraction

GO Biological Process

GO IDTermFold enrichmentFDR

HOMER motifs

MotifFamilyEnrichmentFDR

Enhancers in this class

EnhancerChrLevel-1 clusterCross-support Interaction genesTier 1 pairsOverlap traits

LD blocks

LD blockChrStartEndGenes EnhancersTraitsRowsIn GRBGRB coords

Traits

TraitGenesEnhancersLD blocksOverlap rows Gene-trait setsoverlapEpiGene setsoverlapEpiEnh sets

Candidate evidence

Follow a gene, enhancer or LD block into its genomic context.

GeneSymbolEnhancerTraitLD block Cross-supportTier 1 pairAny-test FDR < 0.05Gene roleLD in GRB

Interactions

Enhancer 1Enhancer 2Interaction scoreBest FDR Permutation FDRRevised RC FDRSignificant testsDirectionCandidate rank

Genomic neighbourhood plot

Genomic neighbourhood

Interactive hg19 view assembled from the complete evidence tables.

Selected gene Qualifying gene Context gene GRB GWAS LD block Selected LD block Enhancer Selected enhancer Tier 1 enhancer

Enter an interval, or select a gene or candidate record.

Glossary and evidence definitions

Terms used in the overlap, pairwise and integrated browser results.

Overlap analysis

GRB
Genomic regulatory block: a conserved long-range regulatory domain containing a predicted target gene and neighbouring bystander genes.
overlapEpiGene
A gene–trait–chromosome set with at least two qualifying, mutually non-overlapping GWAS LD blocks assigned to the gene.
overlapEpiEnh
An overlapEpiGene set in which a significantly associated FANTOM5 enhancer physically overlaps at least one qualifying LD block.
Gene–trait set
One gene, mapped GWAS trait and chromosome considered as a single counting unit.
Overlap row
One enhancer–gene–trait–LD-block association. Several rows may describe the same entity under different traits or blocks.
LD in GRB
The LD-block interval physically overlaps the GRB assigned to the corresponding gene. This differs from the gene merely having a GRB assignment.

Gene roles

GRB-trgt
Predicted long-range regulatory target gene within a GRB.
GRB-byst
Bystander gene located within the target gene’s GRB.
GWAS-trgt / GWAS-byst
GRB target or bystander gene with additional GWAS annotation.
GWAS-out
GWAS-associated gene outside the GRB target and bystander categories.

Pairwise non-additivity

Tier 1 pair
The gene passes the baseline-linear versus pairwise comparison (p < 0.01, ΔBIC < −10), and the pair has permutation FDR < 0.05 with significance in at least three of the four pair-level tests. In Candidate evidence, Tier 1 means that the enhancer participates in such a pair for the displayed gene.
Permutation FDR
BH-adjusted empirical significance of the enhancer product term relative to randomized tissue alignment; this is the strictest individual pair-level test.
Revised RC FDR
BH-adjusted residual-product correlation test after accounting for the baseline-adjusted additive model.
Best FDR
The smallest FDR among the available pair-level tests, including the revised residual-correlation test.
Significant tests
Number of tests significant at FDR < 0.05, including the revised residual-correlation result in the displayed count.
Interaction score
A ranking aid combining −log10(best FDR) with the number of significant tests. It is not an additional hypothesis test.
Direction
Consensus direction of the non-additive signal across compatible tests; it is descriptive rather than a separate significance criterion.
Candidate rank
Ordering within the conservative candidate list; it does not replace the individual statistics.

Integration between analyses

Exact gene–enhancer support
The same gene and enhancer coordinate occur in both the overlap and pairwise analyses.
Cross-support
The strongest correspondence between analyses: exact Tier 1, exact any-test FDR < 0.05, exact tested pair, enhancer-coordinate only, gene only, overlap-only, or pairwise-only.
Enhancer-coordinate only
The enhancer occurs in both analyses, but not with the same gene assignment.
Gene only
The gene occurs in both analyses without an exact shared enhancer coordinate.